The genome sequence of the Whirlpool Ramshorn snail, Anisus vortex (Linnaeus, 1758).

We present a genome assembly from an individual Anisus vortex (the Whirlpool Ramshorn snail; Mollusca; Gastropoda; Hygrophila; Lymnaeoidea; Planorbidae). The genome sequence is 869.5 megabases in span. Most of the assembly is scaffolded into 18 chromosomal pseudomolecules. The mitochondrial genome has also been assembled and is 13.57 kilobases in length.

Revision of the " Chloritisdelibrata (Benson, 1836)" group (Gastropoda, Stylommatophora, Camaenidae).

Chloritisdelibrata (Benson, 1836), known from northeastern India, was believed to have three varietal forms, sometimes mentioned as subspecies: C.delibratavar.khasiensis (Nevill, 1877) and C.delibratavar.fasciata (Godwin-Austen, 1875) from the Khasi Hills, India, and C.delibratavar.procumbens (Gould, 1844) from Dawei in Myanmar. The reproductive anatomy of the latter form is known and does not match with those of any continental camaenid genera, but does with that of the newly examined Chloritisplatytropis Möllendorff, 1894 from Thailand. The latter species is conchologically similar to Bouchetcamaenahuberi Thach, 2018 (synonym of Helixfouresi Morlet, 1886), which is the type species of the genus Bouchetcamaena Thach, 2018. Thus, Bouchetcamaena can provisionally host the entire Chloritisdelibrata -group with the exception of var. fasciata, which is transferred to Burmochloritis Godwin-Austen, 1920 due to the multiple reddish bands on its shell. The examination of shells deposited in the Natural History Museum, London revealed that seven morphologically distinguishable forms are present, which are accepted here as representing distinct species. Four new species are described from India: Bouchetcamaenafoveata Páll-Gergely sp. nov., B.fusca Páll-Gergely sp. nov., B.raripila Páll-Gergely sp. nov., and B.subdelibrata Páll-Gergely sp. nov.

Clarification on the name-bearing type designation of several cyclophorid species (Mollusca, Gastropoda) by H. H. Godwin-Austen (1915).

The type series boundary and the name-bearing type designation of each cyclophorid taxon originally described by Godwin-Austen are clarified based on an interpretation that complies with the ICZN. Previous statuses of type specimens designated by previous authors are reconsidered. Lectotypes of Spiraculum oakesi Godwin-Austen, 1915, Spiraculum kempi Godwin-Austen, 1915, Pterocyclos aborensis Godwin-Austen, 1915, Pterocyclos miriensis Godwin-Austen, 1915, Pterocyclos brahmakundensis Godwin-Austen, 1915, Spiraculum luyorensis Godwin-Austen, 1915, Spiraculum putaoensis Godwin-Austen, 1915, and Theobaldius oakesi Godwin-Austen, 1915 are here designated to stabilize the existing nomenclature. In addition, the type specimens of Pterocyclos miriensis and Theobaldius oakesi are photographed and figured for the first time.

Genus-level revision of the Alycaeidae (Gastropoda, Cyclophoroidea), with an annotated species catalogue.

412 species-group names (including 11 replacement names), and 14 genus-group names of the Alycaeidae have been introduced to date. Type materials of 85% (336) of the known species and subspecies were examined, a further 5% (19) of the taxa were studied using available non-type material, and for another 6% (22) the original descriptions were sufficiently detailed to evaluate their taxonomic status. Only 3% of the taxa (12) could not be examined. Special attention was paid to the sculpture of the embryonic whorls and the sutural tube-microtunnel system in order to provide a novel classification for this group. In this study 363 taxa (320 species or 43 subspecies) are accepted within the family Alycaeidae. Of these, 22 have been described by the lead author and his coauthors in previous publications. In addition, there are 18 species that were formerly classified in Cycloryx and now belong to Pincerna due to its synonymy with Cycloryx. Among the remaining 323 species, 209 (65%) are transferred here to another genus, whilst 114 (35%) have remained in their original genus. Seven genera are accepted. While some questions (e.g., the distinction between Pincerna and Alycaeus) remained unanswered, this revision made three main achievements: (1) The Dicharax species were identified based on the absence of spiral striation on the entire shell; (2) the Metalycaeus species were identified based on the spiral striation of the protoconch; (3) and Stomacosmethis was separated from Alycaeus based on the extremely short sutural tube. Five nominal species are being synonymised with other species, and eight species are now treated as subspecies. The following replacement names are proposed: Dioryx urnula niosiensis Páll-Gergely, nom. nov. for Alycaeus urnula var. daflaensis Godwin-Austen, 1914; Dioryx urnula rotundus Páll-Gergely, nom. nov. for Alycaeus urnula var. globosus Godwin-Austen, 1914; Pincerna crenilabris juttingae Páll-Gergely, nom. nov. for Alycaeus crenilabris laevis van Benthem Jutting, 1959; Pincerna crenilabris korintjiensis Páll-Gergely, nom. nov. for Alycaeus crenilabris latecostatus van Benthem Jutting, 1959; Dicharax conicus jatingaensis Páll-Gergely, nom. nov. for Alycaeus conicus var. nanus Godwin-Austen, 1914; Metalycaeus godwinausteni Páll-Gergely, nom. nov. for Alycaeus neglectus Godwin-Austen, 1914; and finally Metalycaeus suhajdai Páll-Gergely, nom. nov. for Alycaeus varius Godwin-Austen, 1914.

The first global deep-sea stable isotope assessment reveals the unique trophic ecology of Vampire Squid Vampyroteuthis infernalis (Cephalopoda).

Vampyroteuthis infernalis Chun, 1903, is a widely distributed deepwater cephalopod with unique morphology and phylogenetic position. We assessed its habitat and trophic ecology on a global scale via stable isotope analyses of a unique collection of beaks from 104 specimens from the Atlantic, Pacific and Indian Oceans. Cephalopods typically are active predators occupying a high trophic level (TL) and exhibit an ontogenetic increase in δ15N and TL. Our results, presenting the first global comparison for a deep-sea invertebrate, demonstrate that V. infernalis has an ontogenetic decrease in δ15N and TL, coupled with niche broadening. Juveniles are mobile zooplanktivores, while larger Vampyroteuthis are slow-swimming opportunistic consumers and ingest particulate organic matter. Vampyroteuthis infernalis occupies the same TL (3.0-4.3) over its global range and has a unique niche in deep-sea ecosystems. These traits have enabled the success and abundance of this relict species inhabiting the largest ecological realm on the planet.

Notes on the sinistral helicoid snail Bertia cambojiensis (Reeve, 1860) from Vietnam (Eupulmonata, Dyakiidae).

Since the time of the original description there have been no precise locality records in Cambodia of Bertia cambojiensis (Reeve, 1860) and it was believed to be extinct. In 2012, a joint Natural History Museum survey with Vietnamese colleagues rediscovered living populations of this huge sinistral helicoid snail in a protected area of southern Vietnam. The genitalia and radula morphology are re-assessed and type specimens of all recognised congeners are figured herein. The unique morphological characters of this species are a small and simple penis, well-developed amatorial organ complex that incorporates four amatorial organ ducts, a short gametolytic organ complex and spiked papilla, and radula morphology with unicuspid teeth. The type locality of B. cambojiensis, which has been contentious, is determined here to be in the vicinity of 'Brelum', Vietnam, near the border with Cambodia. In addition, the nucleotide sequences of barcoding genes COI, 16SrRNA and 28S fragments were provided for further comparison.

Cryptic diversity of limestone karst inhabiting land snails (Cyclophorus spp.) in northern Vietnam, their evolutionary history and the description of four new species.

Limestone karsts can form terrestrial habitat islands for calcium-dependent organisms. In Vietnam, many karst habitats are threatened, while their rich biodiversity is still far from being thoroughly explored. Given that conservation of karst biota strongly relies on correct species identification, the presence of undetected cryptic species can pose severe problems. The present study focuses on cryptic diversity among karst-inhabiting land snails of the genus Cyclophorus in northern Vietnam, where specimens with a similar shell morphology have been reported from various regions. In order to examine the diversity and evolutionary history of this "widespread morphotype", we generated a Bayesian phylogeny based on DNA sequence data. Automatic Barcode Gap Discovery (ABGD) and the Bayesian implementation of the Poisson tree processes model (bPTP) contributed to species delimitation and analyses of shell shape and size aided the morphological characterisation of individual species. We found that the examined specimens of the widespread morphotype did not form a single monophyletic group in the phylogeny but clustered into several different clades. We delimited nine different species that develop the widespread morphotype and described four of them as new. Processes of convergent evolution were probably involved in the origin of the delimited species, while their generally allopatric distribution could result from interspecific competition. Our findings indicate ongoing processes of speciation and a potential case of morphological character displacement. The high degree of morphological overlap found among the species underlines the importance of DNA sequence data for species delimitation and description in the genus Cyclophorus. Given the findings of the present study and the high potential that as yet undiscovered cryptic taxa have also evolved in other groups of karst-inhabiting organisms, we argue for a systematic and efficient detection and description of Vietnam's karst biodiversity to provide a solid basis for future conservation planning.

An annotated type catalogue of seven genera of operculate land snails (Caenogastropoda, Cyclophoridae) in the Natural History Museum, London.

The collection of the seven cyclophorid snail genera housed in the Natural History Museum, London (NHM), includes 95 available species-level names belonging to the genera Pterocyclos Benson, 1832, Cyclotus Swainson, 1840, Myxostoma Troschel, 1847, Rhiostoma Benson, 1860, Scabrina Blanford, 1863, Crossopoma Martens, 1891, and Pearsonia Kobelt, 1902. Lectotypes are here designated for twelve available species-level names to stabilise existing the nomenclature. A complete catalogue of these types, including colour photographs, is provided for the first time. After examining these type specimens, an unpublished manuscript name was found and is described herein as Pterocyclosanamullayensis Sutcharit & Panha, sp. n.

Type material of land snails (Mollusca: Gastropoda) described from New Zealand by taxonomists in Europe and North America between 1830 and 1934, and the history of research on the New Zealand land snail fauna from 1824 to 1917.

Details are provided on 124 land snail species and varieties from New Zealand, and a further 14 species putatively from New Zealand, all of which were described by European and North American taxonomists between 1830 and 1934, based on specimens collected between 1824 and 1924. Primary type material of 95 of these taxa was located in Northern Hemisphere museums during the present study. Lectotypes are designated for: Helix chimmoi Pfeiffer, 1857, Helix glabriuscula Reeve, 1852, Helix (Paryphanta) gilliesi Smith, 1880, Nanina ? celinde Gray, 1850, Zonites chiron Gray, 1850 and Zonites coma Gray, 1843. Neotypes are designated for Helix conella Pfeiffer, 1861 and Helix tau Pfeiffer, 1861. Primary type material of the following taxa is figured herein for the first time: Bulimus? (Laoma) leimonias Gray, 1850, Cyclophorus cytora Gray, 1850, Cyclostoma (Cyclophorus?) lignarium Pfeiffer, 1857, Helix chimmoi Pfeiffer, 1857, Helix egesta Gray, 1850, Helix fatua Pfeiffer, 1857, Helix greenwoodi Gray, 1850, Helix guttula Pfeiffer, 1853, Helix kermandeci Pfeiffer, 1857, Helix portia Gray, 1850, Helix sciadium Pfeiffer, 1857, Helix venulata Pfeiffer, 1857, Helix (Paryphanta) gilliesi Smith, 1880, Hydrocena (Omphalotropis) vestita Pfeiffer, 1855, Nanina ? celinde Gray, 1850, Nanina erigone Gray, 1850, Nanina mariae Gray, 1843, Patula modicella var. vicinalis Mousson, 1873, Realia egea Gray, 1850, Vitrina kermadecensis Smith, 1873 and Zonites chiron Gray, 1850. New taxonomic combinations introduced herein include: Allodiscus nematophora (Reeve, 1854), Cavellia biconcava (Reeve, 1852), Charopa chimmoi (Pfeiffer, 1857), Coneuplecta regularis (Reeve, 1854), Delos jeffreysiana (Reeve, 1852), Fectola tau (Pfeiffer, 1861), Fectola varicosa (Reeve, 1852), Flammulina crebriflammea (Reeve, 1852), Lyrotropis vestita (Pfeiffer, 1855), ?Neophenacohelix ziczac (Gould, 1846), Parabalea peregrina (Gould, 1847), Phacussa hypopolea (Reeve, 1852), Phenacharopa novoseelandica (Küster, 1852), Phrixgnathus glabriusculus (Reeve, 1852), Phrixgnathus poecilostictus (Reeve, 1852), Thalassohelix obnubila (Reeve, 1852), Tornatellinops novoseelandica (Küster, 1852) and Wainuia urnula (Reeve, 1854). Helix collyrula Reeve, 1852 and Nanina tullia Gray, 1850 are treated as junior synonyms of Phenacohelix (Neophenacohelix) giveni Cumber 1961 nomen protectum and Helix (Huttonella) pseudoleioda Suter, 1890 nomen protectum, respectively. A brief account is given of the history of research on the New Zealand land snail fauna from 1824 to 1917.

An ironic twist of fate: replacement name for Stenogyra gracilenta Morelet, 1885, not Achatina gracilenta Morelet, 1867 (Mollusca, Gastropoda, Achatinidae).

During work on a monograph of Morelet's contributions to malacology (Breure, Audibert Ablett, forthcoming) we found that two of his newly introduced taxa currently have to be considered as homonyms belonging to the genus Subulina H. Beck, 1837 (classified within the family Achatinidae according to Fontanilla et al. (2017: 385)). The older taxon is Achatina gracilenta which was described by Morelet (1867: 79, pl. 7 fig. 2) from [Angola] "Golungo-Alto, au bord du Rio Quiapose, près de Sange; les environs de Lopollo (district de Huilla)". According to Naggs (1990a: 31; also pers. comm. 10 October 2017) this species has to be classified as Subulina gracilenta (Morelet, 1867). The type material of this species is present in the Natural History Museum, London (NHMUK 1893.2.4.263-265). The younger taxon was described by Morelet (1885: 25, pl. 2 fig. 8) as Stenogyra gracilenta from [Gabon] "environs de Mayumba". Pilsbry (1906: 82) already recognised this as Subulina gracilenta (Morelet, 1885); Naggs (pers. comm. 10 October 2017) confirmed this classification and concluded that this taxon is a junior secondary homonym of Achatina gracilenta Morelet, 1867. Type material for Stenogyra gracilenta Morelet, 1885 has not been located. We here propose Subulina bruggenorum nom. nov. as a replacement name for Stenogyra gracilenta Morelet, 1885 (not Achatina gracilenta Morelet, 1867).

Competition matters: Determining the drivers of land snail community assembly among limestone karst areas in northern Vietnam.

The insular limestone karsts of northern Vietnam harbor a very rich biodiversity. Many taxa are strongly associated with these environments, and individual species communities can differ considerably among karst areas. The exact processes that have shaped the biotic composition of these habitats, however, remain largely unknown. In this study, the role of two major processes for the assembly of snail communities on limestone karsts was investigated, interspecific competition and filtering of taxa due to geographical factors. Communities of operculate land snails of the genus Cyclophorus were studied using the dry and fluid-preserved specimen collections of the Natural History Museum, London. Phylogenetic distances (based on a Bayesian analysis using DNA sequence data) and shell characters (based on 200 semilandmarks) were used as proxies for ecological similarity and were analyzed to reveal patterns of overdispersion (indicating competition) or clustering (indicating filtering) in observed communities compared to random communities. Among the seven studied karst areas, a total of 15 Cyclophorus lineages were found. Unique communities were present in each area. The analyses revealed phylogenetic overdispersion in six and morphological overdispersion in four of seven karst areas. The pattern of frequent phylogenetic overdispersion indicated that competition among lineages is the major process shaping the Cyclophorus communities studied. The Coastal Area, which was phylogenetically overdispersed, showed a clear morphological clustering, which could have been caused by similar ecological adaptations among taxa in this environment. Only the community in the Cuc Phuong Area showed a pattern of phylogenetic clustering, which was partly caused by an absence of a certain, phylogenetically very distinct group in this region. Filtering due to geographical factors could have been involved here. This study shows how museum collections can be used to examine community assembly and contributes to the understanding of the processes that have shaped karst communities in Vietnam.

Illustrated type catalogue of Amphidromus Albers, 1850 in the Natural History Museum, London, and descriptions of two new species.

The collection of the Southeast Asian tree snail genus Amphidromus Albers, 1850 at the Natural History Museum, London includes more than 100 lots of type specimens representing 85 name-bearing types, 9 paratypes and 6 paralectotypes, and one nomen nudum. Lectotypes are here designated for Amphidromuscambojiensis, Amphidromusperakensisglobosus, Amphidromuscolumellarisgloriosa, Amphidromusmaculiferusinflata, Amphidromuslepidus, Amphidromussinistralislutea, Amphidromusmoniliferus, Amphidromusmaculiferusobscura, Amphidromussinistralisrosea and Amphidromussinensivicaria. In addition, the missing types of A.A. Gould were discovered and their type status is discussed. A complete catalogue of these types, including colour photographs is provided for the first time. After examining these type specimens, two new Amphidromus species, Amphidromus (Syndromus) globonevilli Sutcharit & Panha, sp. n. and Amphidromus (Syndromus) principalis Sutcharit & Panha, sp. n. were recognized and are described herein.

Annotated type catalogue of the Megaspiridae, Orthalicidae, and Simpulopsidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London.

The type status is described for 65 taxa of the Orthalicoidea, classified within the families Megaspiridae (14), Orthalicidae (30), and Simpulopsidae (20); one taxon is considered a nomen inquirendum. Lectotypes are designated for the following taxa: Helixbrephoides d'Orbigny, 1835; Simpulopsiscumingi Pfeiffer, 1861; Bulimulus (Protoglyptus) dejectus Fulton, 1907; Bulimusiris Pfeiffer, 1853. The type status of Bulimussalteri Sowerby III, 1890, and Strophocheilus (Eurytus) subirroratus da Costa, 1898 is now changed to lectotype according Art. 74.6 ICZN. The taxa Bulimusloxostomus Pfeiffer, 1853, Bulimusmarmatensis Pfeiffer, 1855, Bulimusmeobambensis Pfeiffer, 1855, and Orthalicuspowissianusvar.niveusPreston 1909 are now figured for the first time. The following taxa are now considered junior subjective synonyms: Bulimusmarmatensis Pfeiffer, 1855 = Helix (Cochlogena) citrinovitrea Moricand, 1836; Vermiculatus Breure, 1978 = Bocourtia Rochebrune, 1882. New combinations are: Kuschelenia (Bocourtia) Rochebrune, 1882; Kuschelenia (Bocourtia) aequatoria (Pfeiffer, 1853); Kuschelenia (Bocourtia) anthisanensis (Pfeiffer, 1853); Kuschelenia (Bocourtia) aquila (Reeve, 1848); Kuschelenia (Bocourtia) badia (Sowerby I, 1835); Kuschelenia (Bocourtia) bicolor (Sowerby I, 1835); Kuschelenia (Bocourtia) caliginosa (Reeve, 1849); Kuschelenia (Bocourtia) coagulata (Reeve, 1849); Kuschelenia (Bocourtia) cotopaxiensis (Pfeiffer, 1853); Kuschelenia (Bocourtia) filaris (Pfeiffer, 1853); Karaindentata (da Costa, 1901); Clathrorthalicusmagnificus (Pfeiffer, 1848); Simpulopsis (Eudioptus) marmartensis (Pfeiffer, 1855); Kuschelenia (Bocourtia) nucina (Reeve, 1850); Kuschelenia (Bocourtia) ochracea (Morelet, 1863); Kuschelenia (Bocourtia) peaki (Breure, 1978); Kuschelenia (Bocourtia) petiti (Pfeiffer, 1846); Clathrorthalicusphoebus (Pfeiffer, 1863); Kuschelenia (Bocourtia) polymorpha (d'Orbigny, 1835); Scholvieniaporphyria (Pfeiffer, 1847); Kuschelenia (Bocourtia) purpurata (Reeve, 1849); Kuschelenia (Bocourtia) quechuarum Crawford, 1939; Quechuasalteri (Sowerby III, 1890); Kuschelenia (Bocourtia) subfasciata Pfeiffer, 1853; Clathrorthalicusvictor (Pfeiffer, 1854). In an addedum a lectotype is being designated for Bulimulus (Drymaeus) interruptusvar.pallidus Preston, 1909. An index is included to all taxa mentioned in this paper and the preceding ones in this series (Breure and Ablett 2011, 2012, 2014).

Systematics of the family Plectopylidae in Vietnam with additional information on Chinese taxa (Gastropoda, Pulmonata, Stylommatophora).

Vietnamese species from the family Plectopylidae are revised based on the type specimens of all known taxa, more than 600 historical non-type museum lots, and almost 200 newly-collected samples. Altogether more than 7000 specimens were investigated. The revision has revealed that species diversity of the Vietnamese Plectopylidae was previously overestimated. Overall, thirteen species names (anterides Gude, 1909, bavayi Gude, 1901, congesta Gude, 1898, fallax Gude, 1909, gouldingi Gude, 1909, hirsuta Möllendorff, 1901, jovia Mabille, 1887, moellendorffi Gude, 1901, persimilis Gude, 1901, pilsbryana Gude, 1901, soror Gude, 1908, tenuis Gude, 1901, verecunda Gude, 1909) were synonymised with other species. In addition to these, Gudeodiscushemmeni sp. n. and Gudeodiscusmessageriraheemi ssp. n. are described from north-western Vietnam. Sixteen species and two subspecies are recognized from Vietnam. The reproductive anatomy of eight taxa is described. Based on anatomical information, Halongella gen. n. is erected to include Plectopylisschlumbergeri and Plectopylisfruhstorferi. Additionally, the genus Gudeodiscus is subdivided into two subgenera (Gudeodiscus and Veludiscus subgen. n.) on the basis of the morphology of the reproductive anatomy and the radula. The Chinese Gudeodiscusphlyariuswerneri Páll-Gergely, 2013 is moved to synonymy of Gudeodiscusphlyarius. A spermatophore was found in the organ situated next to the gametolytic sac in one specimen. This suggests that this organ in the Plectopylidae is a diverticulum. Statistically significant evidence is presented for the presence of calcareous hook-like granules inside the penis being associated with the absence of embryos in the uterus in four genera. This suggests that these probably play a role in mating periods before disappearing when embryos develop. Sicradiscusmansuyi is reported from China for the first time.

An annotated catalogue of type specimens of the land snail genus Cyclophorus Monfort, 1810 (Caenogastropoda, Cyclophoridae) in the Natural History Museum, London.

The collection of land caenogastropod snails in the genus Cyclophorus Monfort, 1810 housed in the Natural History Museum, London (NHM), includes 52 type lots. Lectotypes have been designated for 43 available species-level names to stabilize existing nomenclature, two previously designated lectotype, two holotypes, one paratype, one syntype, one possible syntype and two paralectotypes are also listed. A complete catalogue of the Cyclophorus types in NHM, London is provided for the first time.

Annotated type catalogue of the Bulimulidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London.

The type status is described of 404 taxa classified within the family Bulimulidae (superfamily Orthalicoidea) and kept in the London museum. Lectotypes are designated for Bulimus aurifluus Pfeiffer, 1857; Otostomus bartletti H. Adams, 1867; Helix cactorum d'Orbigny, 1835; Bulimus caliginosus Reeve, 1849; Bulimus chemnitzioides Forbes, 1850; Bulimus cinereus Reeve, 1849; Helix cora d'Orbigny, 1835; Bulimus fallax Pfeiffer, 1853; Bulimus felix Pfeiffer, 1862; Bulimus fontainii d'Orbigny, 1838; Bulimus fourmiersi d'Orbigny, 1837; Bulimus (Mesembrinus) gealei H. Adams, 1867; Bulimus gruneri Pfeiffer, 1846; Bulimus humboldtii Reeve, 1849; Helix hygrohylaea d'Orbigny, 1835; Bulimus jussieui Pfeiffer, 1846; Bulimulus (Drymaeus) binominis lascellianus E.A. Smith, 1895; Helix lichnorum d'Orbigny, 1835; Bulimulus (Drymaeus) lucidus da Costa, 1898; Bulimus luridus Pfeiffer, 1863; Bulimus meleagris Pfeiffer, 1853; Bulimus monachus Pfeiffer, 1857; Bulimus montagnei d'Orbigny, 1837; Helix montivaga d'Orbigny, 1835; Bulimus muliebris Reeve, 1849; Bulimus nigrofasciatus Pfeiffer in Philippi 1846; Bulimus nitelinus Reeve, 1849; Helix oreades d'Orbigny, 1835; Helix polymorpha d'Orbigny, 1835; Bulimus praetextus Reeve, 1849; Bulinus proteus Broderip, 1832; Bulimus rusticellus Morelet, 1860; Helix sporadica d'Orbigny, 1835; Bulimus sulphureus Pfeiffer, 1857; Helix thamnoica var. marmorata d'Orbigny, 1835; Bulinus translucens Broderip in Broderip and Sowerby I 1832; Helix trichoda d'Orbigny, 1835; Bulinus ustulatus Sowerby I, 1833; Bulimus voithianus Pfeiffer, 1847; Bulimus yungasensis d'Orbigny, 1837. The type status of the following taxa is changed to lectotype in accordance with Art. 74.6 ICZN: Bulimulus (Drymaeus) caucaensis da Costa, 1898; Drymaeus exoticus da Costa, 1901; Bulimulus (Drymaeus) hidalgoi da Costa, 1898; Bulimulus (Drymaeus) interruptus Preston, 1909; Bulimulus (Drymaeus) inusitatus Fulton, 1900; Bulimulus latecolumellaris Preston, 1909; Bulimus (Otostomus) napo Angas, 1878; Drymaeus notabilis da Costa, 1906; Drymaeus notatus da Costa, 1906; Bulimulus (Drymaeus) nubilus Preston, 1903; Drymaeus obliquistriatus da Costa, 1901; Bulimus (Drymaeus) ochrocheilus E.A. Smith, 1877; Bulimus (Drymaeus) orthostoma E.A. Smith, 1877; Drymaeus expansus perenensis da Costa, 1901; Bulimulus pergracilis Rolle, 1904; Bulimulus (Drymaeus) plicatoliratus da Costa, 1898; Drymaeus prestoni da Costa, 1906; Drymaeus punctatus da Costa, 1907; Bulimus (Leptomerus) sanctaeluciae E.A. Smith, 1889; Bulimulus (Drymaeus) selli Preston, 1909; Drymaeus subventricosus da Costa, 1901; Bulimulus (Drymaeus) tigrinus da Costa, 1898; Drymaeus volsus Fulton, 1907; Drymaeus wintlei Finch, 1929; Bulimus zhorquinensis Angas, 1879; Bulimulus (Drymaeus) ziczac da Costa, 1898. THE FOLLOWING JUNIOR SUBJECTIVE SYNONYMS ARE ESTABLISHED: Bulimus antioquensis Pfeiffer, 1855 = Bulimus baranguillanus Pfeiffer, 1853; Drymaeus bellus da Costa, 1906 = Drymaeus blandi Pilsbry, 1897; Bulimus hachensis Reeve 1850 = Bulimus gruneri Pfeiffer, 1846 = Bulimus columbianus Lea, 1838; Bulimus (Otostomus) lamas Higgins 1868 = Bulimus trujillensis Philippi, 1867; Bulimulus (Drymaeus) binominis lascellianus E.A. Smith, 1895 = Bulimulus (Drymaeus) binominis E.A. Smith, 1895; Drymaeus multispira da Costa, 1904 = Helix torallyi d'Orbigny, 1835; Bulimulus (Drymaeus) plicatoliratus Da Costa, 1898 = Bulimus convexus Pfeiffer, 1855; Bulimus sugillatus Pfeiffer, 1857 = Bulimus rivasii d'Orbigny, 1837; Bulimus meridionalis Reeve 1848 [June] = Bulimus voithianus Pfeiffer, 1847. NEW COMBINATIONS ARE: Bostryx montagnei (d'Orbigny, 1837); Bostryx obliquiportus (da Costa, 1901); Bulimulus heloicus (d'Orbigny, 1835); Drymaeus (Drymaeus) lusorius (Pfeiffer, 1855); Drymaeus (Drymaeus) trigonostomus (Jonas, 1844); Drymaeus (Drymaeus) wintlei Finch, 1929; Drymaeus (Mesembrinus) conicus da Costa, 1907; Kuschelenia (Kuschelenia) culminea culminea (d'Orbigny, 1835); Kuschelenia (Kuschelenia) culmineus edwardsi (Morelet, 1863); Kuschelenia (K.) gayi (Pfeiffer, 1857); Kuschelenia (Kuschelenia) tupacii (d'Orbigny, 1835); Kuschelenia (Vermiculatus) anthisanensis (Pfeiffer, 1853); Kuschelenia (Vermiculatus) aquilus (Reeve, 1848); Kuschelenia (Vermiculatus) bicolor (Sowerby I, 1835); Kuschelenia (Vermiculatus) caliginosus (Reeve, 1849); Kuschelenia (Vermiculatus) cotopaxiensis (Pfeiffer, 1853); Kuschelenia (Vermiculatus) filaris (Pfeiffer, 1853); Kuschelenia (Vermiculatus) ochracea (Morelet, 1863); Kuschelenia (Vermiculatus) petiti (Pfeiffer, 1846); Kuschelenia (Vermiculatus) purpuratus (Reeve, 1849); Kuschelenia (Vermiculatus) quechuarum (Crawford, 1939); Naesiotus cinereus (Reeve, 1849); Naesiotus dentritis (Morelet, 1863); Naesiotus fontainii (d'Orbigny, 1838); Naesiotus orbignyi (Pfeiffer, 1846); Protoglyptus pilosus (Guppy, 1871); Protoglyptus sanctaeluciae (E.A. Smith, 1889). TYPE MATERIAL OF THE FOLLOWING TAXA IS FIGURED HEREIN FOR THE FIRST TIME: Bulimus cinereus Reeve, 1849; Bulimus coriaceus Pfeiffer, 1857; Bulimulus laxostylus Rolle, 1904; Bulimus pliculatus Pfeiffer, 1857; Bulimus simpliculus Pfeiffer, 1855.

Annotated type catalogue of the Bothriembryontidae and Odontostomidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London.

The type status is described for specimens of 84 taxa classified within the families Bothriembryontidae and Odontostomidae (superfamily Orthalicoidea) and kept in the Natural History Museum, London. Lectotypes are designated for Bulimus (Liparus) brazieri Angas, 1871; Bulimus broderipii Sowerby I, 1832; Bulimus fuligineus Pfeiffer, 1853; Helix guarani d'Orbigny, 1835; Bulimus (Tomigerus) ramagei E.A. Smith, 1890; Helix rhodinostoma d'Orbigny, 1835; Bulimus (Bulimulus) ridleyi E.A. Smith, 1890. The type status of the following taxa is changed to lectotype in accordance with Art. 74.6 ICZN: Placostylus (Euplacostylus) cylindricus Fulton, 1907; Bulimus pyrostomus Pfeiffer, 1860; Bulimus turneri Pfeiffer, 1860. The following taxon is synonymised: Bulimus oblitus Reeve, 1848 = Bahiensis neglectus (Pfeiffer, 1847).

Annotated type catalogue of the Amphibulimidae (Mollusca, Gastropoda, Orthalicoidea) in the Natural History Museum, London.

The type status is described of 39 taxa classified within the family Amphibulimidae (superfamily Orthalicoidea) and kept in the London museum. One taxon, Bulimus elaeodes Pfeiffer, 1853, is removed to the Strophocheilidae. Lectotypes are designated for Bulimus adoptus Reeve, 1849; Bulimus (Eurytus) eros Angas, 1878; Helix onca d'Orbigny, 1835; Amphibulima pardalina Guppy, 1868. The type status of the following taxon is changed to lectotype in accordance with Art. 74.6 ICZN: Strophocheilus (Dryptus) jubeus Fulton, 1908.As general introduction to this and following papers on Orthalicoid types in the Natural History Museum, a brief history of the London collection is given and several examples of handwriting from different authors are presented.